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Table 2 Semantic roles

From: Construction of an annotated corpus to support biomedical information extraction



EXAMPLE (bold = semantic argument, italics = event central verb)


Drives/instigates event

The narL gene product activates the nitrate reductase operon


a) Affected by/results from event

b) Focus of events describing states

recA protein was induced by UV radiation

The ptsH mutant lacks HPr


Method/way in which event is carried out

cpxA gene increases the levels of csgA transcription by dephosphorylation of CpxR


Used to carry out event

EnvZ functions through OmpR to control NP porin gene expression in E. coli.


Where complete event takes place

Phosphorylation of OmpR modulates expression of the ompF and ompC genes in Escherichia coli


Start point of event

A transducing lambda phage was isolated from a strain harboring a glpD"lacZ fusion


End point of event

Transcription is activated by binding of the cyclic AMP (cAMP)-cAMP receptor protein (CRP) complex to a CRP binding site


Situates event in time/w.r.t another event

The Alp protease activity is detected in cells after introduction of plasmids


Environmental conditions/changes in conditions

Strains carrying a mutation in the crp structural gene fail to repress ODC and ADC activities in response to increased cAMP


Change of level or rate

marR mutations elevated inaA expression by 10- to 20-fold over that of the wild-type.


Descriptive information about AGENT of event

HyfR acts as a formate-dependent regulator


Descriptive information about THEME of event

The ptsH mutant lacks HPr.


Purpose/reason for the event occurring

The fusion strains were used to study the regulation of the cysB gene

  1. For each semantic role, a brief description is given, together with an example sentence containing an instance of the role. In the example sentences, the verb on which the event is centred is indicated in italics, whilst the event argument corresponding to the appropriate semantic role is indicated in bold.