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Table 2 Example calculation of Position Shift Error (PSE). See Equation: 4 and associated text for explanation.

From: OXBench: A benchmark for evaluation of protein multiple sequence alignment accuracy

a) Reference Alignment

1

2

3

3.5

4

5

6

7

G

K

L

-

M

R

A

Y

G

K

P

H

K

L

R

L

1

2

3

4

5

6

7

8

b) Test Alignment

1

2

3

4

4.5

5

6

7

G

K

L

M

-

R

A

Y

G

K

P

H

K

L

R

L

1

2

3

4

5

6

7

8